Never one to give up, Optimus managed to down Devastator in return, only to find himself at Megatron's mercy. The Decepticon leader pointed out to him the humans' self-obsessed behavior in the face of the devastation, suggesting the fleshlings were worthless, and the Autobots should join the Decepticons in wiping them out. Prime Directive #5 The heroic actions of a team of firefighters and other humans proved to Optimus that the humans were worth saving, and the Autobots delivered a smackdown on the Decepticons. The fight was interrupted by the news that a nuclear missile was inbound, and despite Prime's protests, Superion sacrificed himself intercepting it. Prime recovered from the shock wave to find that the Decepticons had fled. He appealed to Grimlock to return to the Autobot ranks, an offer turned down by the Dinobot. Prime Directive #6
Transformers: Cybertron, named Galaxy Force in Japan, was a follow-up to the successful Armada and Energon lines. The companion anime series program was not originally conceived or produced as a continuation of the Armada/Energon storyline, and it was only later that it was retconned to be a continuation. Thus, several inconsistencies exist. Once again, a new feature was introduced for the line—Cyber Keys, which activated special functions on the figures in a manner similar to the Mini-Cons in Armada. Each figure came with a Cyber Key regardless of size, but there were several Cyber Keys crucial to the plot of the anime series which only came with certain figures. Again, many of the same characters were included but with entirely new figures from new molds.

This Trailer is identical to the original BeCool Trailer other than the stickers, and now comes with the trailer/launcher accessory from Bot Shots redecoed in blue with red stripes. While the vehicle mode sticker in his chest remains, the hand-holding and sleeping ones have been replaced by pictures of the launcher: one in base mode and one in trailer mode. He also comes with a sticker sheet for decorating the Trailer Base.
Like the previous Prime, it is loaded with features, though it ditches the first movie's Automorphing for the new Mech Alive gimmick. When a grey tab on his midriff is pressed, Prime's entire pectoral region "flexes", lights in his chest and eyes glow, and he declares "I am Optimus Prime" in Peter Cullen's voice. For the European release of the figure, to get past the language barrier, the Cullen sound clip is abbreviated to just "Optimus Prime"; in some regions, including Quebec and the UK (but only after a month or two), both versions were made available. A transformation sound effect activates when his torso is transformed, and once Prime is in truck mode, an almost unnoticeable button (a small square located beside the screw for the battery compartment, undocumented in his instructions) is revealed, which activates engine-idling noises.
Part of the first wave of Bot Shots Series 1 single-packs, Optimus Prime is a teeny little long-nosed semi truck with "trailer", with a spring-loaded automatic transformation to robot mode triggered when his front bumper is pressed. He has a "spinner" in his chest that shows his three attack types and power levels. His truck mode and robot body are influenced by the live-action Optimus Prime, while his head is taken from the original. His blue paint is light and sparkly-metallic, his red paint is dull matte, and his optics are yellow.
Optimus was eventually revived by the Autobots when they arrived to liberate Master from Decepticon hands. They found allies in the native Cyberdroids—those Optimus loyalists who had fought the Malignus insurgency. Autobots and Cyberdroids liberated the deposed triumvirate of Apex, Hi-Q, and Diac. Hoping to check the power-mad Galvatron and his three Cyberdroid allies, the Autobot leader was revived and binary-bonded himself to three Cyberdroids, including Surge, and Apex, who formed his head. Thus empowered, "Triple-Threat Prime" battled Galvatron and his forces across Master in a series of epic conflicts. The Autobots were unsuccessful and were forced to retreat. A Brush With Infamy–Prologue
A retool of Crash Combiner Optimus Prime available in a two-pack with Grimlock, featuring a new Optimus Prime-ish Combiner head and new arm parts with a green dinosaur-themed front shoulder pads, and he features a lighter shade of red plastic. Like the previous versions, Optimus Prime can become the top half of any Crash Combiner gestalt, though his official combination is with Grimlock as Primelock.
In reorganizing the company, LEGO became more efficient. The data shows that they have been able to standardize costs and it is likely that they were able to hedge the price of plastic against future price fluctuations. In protecting their business, they have also been able to normalize prices for their customers. A stable price is good for everyone; it helps LEGO ensure that future production costs are budgeted for and it helps the consumer manage their budget. This allows LEGO to protect and grow its market share.

Released in the latter stages of the Cybertron toyline, this redeco of the Leader-class Optimus Prime figure introduced a new colour scheme that several other tail-end releases would emulate. Taking its name from the Japanese version of Cybertron and with a bio noting that this new body depicts Prime after he embarks on his galaxy-spanning mission at the end of the Cybertron cartoon, "Galaxy Force" Optimus Prime replaces the original toy's blue parts with black and the translucent blue with smokey clear, and moulds several—but not all—grey parts out of red plastic, including the toy's rifles, rail gun covers, missile launchers, and Super Mode leg guns. In the realm of new paint operations, Prime now sports all four Cyber Planet Key symbols on his shoulders, and his unique Matrix-shaped Cyber Key (now with the code lo9x) finally has a gold-painted border, as it had always had in Japan. Additionally, Prime's hip joints were tightened up with this release, making it easier to stand him up in Super Mode.
In robot mode, like Combat Hero Prime, the bottom of the truck became his familiar windowed chest based upon the original Optimus Prime toy. His main gimmick was light up LED headlights, and a lightpipe in the bottom of his right fist, illuminated by an LED in his right forearm, that would illuminate his clear sword or his double-barreled rifle if they were used in that hand. All three LEDs were activated simultaneously by pressing the sunroof-like depression on the roof of the truck.
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