Optimus was one of the many soldiers fighting under the command of the Autobot leader, Jetfire. He was among the Autobots who launched in the Ark to clear a path through an asteroid belt. When the Decepticons attacked the ship, he fought a Seeker Decepticon while Jetfire sent the Ark crashing into Earth, knocking everyone aboard offline. Four million years later, they were repaired when the ship's systems were reactivated by a volcanic eruption. In the Transformers
The TakaraTomy "Autobot Alliance" release of this toy has some significant differences, including casting the red plastic in a darker shade, making his shin and wheel joints grey rather than Hasbro's bronze, painting the Matrix of Leadership silver, and swapping out the dual energon swords for battle hooks, similar to 2010's Leader class Optimus Prime. However, due to the Japanese release retaining the Hasbro packaging, the box still advertises the dual energon swords rather than the hooks. He retains the masked face used for the Hasbro release.