Studio Series Optimus Prime is an all new Voyager Class mold that transforms from robot into a Peterbilt 379 semi-truck. He comes with a pair of Energon Swords, which can be mounted onto his arms or store on the back of his robot mode or truck mode. The figure combines engineering concepts from his Age of Extinction Voyager Class figure with techniques previously used on Prime: First Edition Optimus Prime. He also features a more subdued color scheme compared to other Optimus Prime figures. Like the rest of the Studio Series figures, the figure prioritizes robot mode scale and thus stands shorter than his Voyager Class wavemate Starscream. The CG renders depict him possessing long smokestacks, but the final product has them shortened... again. He also comes with a cardboard backdrop display depicting the climatic forest battle from Revenge of the Fallen.
Similar to the Activators toyline, One-Step Changer Optimus Prime features a spring-loaded transformation. He features swivel articulation in the head and both arms. He shares a similar (not identical) transformation with his fellow One-Step changer, Hound. Also, due to his transformation scheme, his shoulders can be pushed forward (the arms is pushed on the same time), which also allows him to wield a close ranged weapons with longer handles with both hands.
As the Binaltech saga unfolds, the multiverse is threatened with destruction, until the original continuity is restored and the Binaltech universe is split into and preserved within its own separate universe. Overdrive confirms that the threat is over by visiting the dimension occupied by the original timeline, and reveals that in that world, Hot Rod had received the Matrix and taken over leadership of the Autobots as Rodimus Prime.
When Sideswipe was accused and convicted of destroying the Nova Suspension Bridge, Optimus was infuriated that one of his own Autobots could commit such a heinous act. Optimus was tasked to hunt down and deactivate Sideswipe, though the task proved to be easy as Sideswipe did not run, hide or resist. After Optimus had deactivated Sideswipe, he was approached by Prowl, who pointed out that Sideswipe's actions made little sense, as he had committed the crime in broad daylight before many witnesses. Agreeing that the Decepticons might be behind the situation, Optimus allowed Prowl to take a team of Autobots to Carob Island to clear Sideswipe's name while Optimus and Jazz stayed behind to continue working on human/Autobot relations. Prowl returned with evidence that it was indeed a Decepticon-controlled clone of Sideswipe that had committed the crime. Optimus then presumably reactivated Sideswipe when his name was cleared. Deadly Paradise