This figure was only available in a two-pack with a Voyager Class Grimlock figure. The two-pack was exclusively available at Target stores in the United States, Canada and Australia, at Tesco stores in the United Kingdom and Hungary and at Galeria Kaufhof department stores in Germany. In Asia, most of the remaining stock are available in some hobby shops and some video stores like Speedy Video.
In his first appearance in the series, Optimus looks similar to the original G1 Optimus Prime until he copies himself with a semi-truck and his design in robot mode changes to the image shown here. Soon after, he retrieves his Mini-Con partner, Sparkplug. The cab of his truck mode detaches to form Optimus himself, while the trailer transforms into a battle station mode operable by Optimus and several Mini-Cons. Optimus can also combine with his base to form a "Super Mode" robot that, when combined with Sparkplug, can fire a series of powerful lasers. Additionally, Optimus can combine with Jetfire and/or Overload in his Super Mode for additional power.
Generation 2 Optimus Prime comes with all his original accessories, as well as two major new play features. To increase his arsenal, he is armed with a pair of black missile launchers that fit in his fists and fire red spring-loaded missiles; these launchers were retooled from the G.I. Joe figure Barricade. Secondly, he comes with an electronic "sound and lights module" that can either plug into the front of the trailer (where it resembles the air conditioners used on refrigerated trailers), or hook onto Optimus's back in robot mode (via a new slot sculpted into the cab robot). Unfortunately, the soundbox is heavy enough that doing so can easily overpower Prime's hip joints, causing him to topple backward. The soundbox features a large, vacuum metalized Autobot symbol on its front, and three grey buttons which trigger two flashing red LEDs on either side of the module, and a trio of sound effects: a truck engine noise, a laser gun sound effect and a garbled voice that said "I am Optimus Prime!". The module also features ports on either side into which the missile launchers can peg.
Released as part of the "W Convoy" campaign ("Double" Convoy, that is!) alongside a silver chrome version of Energon Rodimus, this gold chrome "Mēki" ("plating") version of the Deluxe size Optimus Prime toy was available only as a prize in a mail-in Robot Points contest. Initially limited to 20 units in the original draw, it appears that many more were made available through toy store drawings, given how many went on to appear on online auction sites through 2005 and 2006.
If Sparkplug suggested that the Autobots not worry about subtlety, the entire squad transformed and began prowling the streets in search of Decepticons. Oblivious to the panic they were causing, Optimus assumed that the Decepticons were spotted nearby when a police officer declared that the city was under siege by giant robots. Unable to restore order in time to halt the Decepticons’ plot, Optimus and the rest of his team were destroyed in the ensuing earthquake.
The back story of the Transformers on ancient Cybertron is told in Transformers: Defiance. In this story, Starscream returns to Cybertron after encountering an enemy scout ship in the Eshems Nebula. After being repaired for minor damage by Ratchet, Starscream and Ironhide report to Optimus Prime and Megatron. In issue #2, Optimus Prime is ordered by Megatron to counter-attack Cybertron's invaders, but instead, Optimus organizes a resistance to Megatron's decision. In issue #3, Optimus Prime, Jazz, and Prowl break into Megatron's quarters while he is away looking for clues to why he is acting so strangely. What they find is an ancient artifact that Megatron had restored.
Part of the first wave of TakaraTomy Lost Age: Movie Advanced toys, this Japanese-exclusive redeco of 2007 Transformers Protoform Optimus Prime is given a Generation 1-styled color scheme (which is also based on his appearances in IDW's Transformers: Foundation comic-book mini-series). Protoform Optimus Prime transforms into his "entry" mode. This mode somewhat resembles a cybertronic truck, with false wheels sculpted onto the sides, and Optimus Prime's trademark windows in front. His now-blue flame decoration piece fits to the back of the "vehicle", creating the illusion of a flaming comet's tail (Not sure if this was mean to represent his jetpack blast effect or his atmospheric entry heat effect, which is for no reason: entirely blue instead of hot red). The bottom of the vehicle has small wheels to roll the toy across smooth surfaces.