He does not come with his Mini-Con partner Over-Run in this release; instead, Optimus Prime and his wave-mate Demolishor debuted a new brand of Cyber Key: retooled from the standard Cybertron style key, this version lacked a sculpted insignia, so that any symbol could be tampographed on to it, and featured a Mini-Con-style Powerlinx socket on the back, allowing it to operate the figure's formerly-Mini-Con-activated "punching" feature without the figure itself having to be retooled.
Laser Optimus Prime transforms into a mid-1990s Western Star 4964EX, pulling a grey and black fuel tank trailer. The sides of the trailer are decorated with a sticker depicting Prime using a flamethrower to burn down a banner with the words "Optimus Prime Octane" on it. The truck is armed with a disc launcher on the roof of its trailer, which launches five blue discs bearing the Generation 2 Autobot insignia as a knob is turned. Like the other "Laser" toys released this year, Prime features electronics powered by two button-cell batteries; in truck mode, pressing the button on the cab's roof causes his headlight to light up with yellow LEDs.
While stock photos and renders show the figure with a faceplate, this image posted by the Transformers Facebook page shows him with his mouth exposed. This was later found as a variant at least once. A running change was made, changing the blades to be more translucent. The semi-translucent blades release features a date code "73461", whereas the translucent orange blades version features a date code written as "80171".
In The Transformers: Devastation, Hot Rod is repaired and reformatted into a Dodge Viper. Dealer tried to convince him to pull some strings to get him to Earth, but Hot Rod and Wheeljack had the more important task of rescuing Ironhide. They ran into a bit of trouble with the Machination's Headmasters. They managed to escape temporarily, but were found and Wheeljack was incapacitated. Hot Rod easily held his own with the Headmasters, and noted that for some reason they had lost their "edge". He received a little help from Hardhead, who was mainly there to take Hot Rod and the others back to the Ark-32 for evacuation to Garrus-9. Hot Rod refused to go, wanting to see the Headmaster mystery through to the end. Hardhead reluctantly let him go.
Early in this new series of encounters with Megatron, Red Alert contacted the Autobots on Earth and asked for Optimus Prime to join then on Velocitron. Ransack and Crumplezone tricked Hot Shot and Dirt Boss into believing that a race between them would determine which of them would be allowed to race Override for the Planet Cup. During the race, Ransack and Crumplezone caused a rockslide that buried Hot Shot. Hot shot was saved when Optimus Prime, Vector Prime, and Landmine arrived, but Optimus was upset that Hot Shot had become involved with the locals. Clocker and Brakedown were eager to help the Autobots. After being unable to obtain the Planet Cup from Override herself, Optimus Prime endorsed Hot Shot's plan to win the cup in a race with the aide of the other Autobots. Ransack and Crumplezone were contacted by Megatron and were told to keep on eye on things. They also decided to enter the race, either to win the cup and cause as much damage as they could to the other racers. Override, Dirt Boss, Ransack, Crumplezone, Optimus Prime, Hot Shot, Red Alert, Landmine, Clocker and Brakedown participated in the qualification round.
Entertainment Pack Optimus Prime is a slight redeco of the SDCC figure, with some deco differences between the SDCC release; this version has gray windows instead of blue and is missing some of the silver deco on the helmet and roof/feet, as well as the yellow paint on his lights. His midriff is also darker. Optimus also comes with a similarly-scaled Megatron and PVC figurines of Jack Darby, Miko Nakadai, and Raf Esquivel.
The Transformers Energon figure pen is based on Optimus Prime in his standard Energon body. About the size of a scout class toy, he transforms from truck mode to robot mode almost identically to his mainline toys, apart from the fact that his feet contain the pen and cannot bend. It's really hard to write with, and almost impossible to pose, even though it has full arm articulation. Amusingly, this figure can position itself to transform into his super robot mode (by flipping his helmet on, opening his chest and flipping his hands in), but because there are no versions of the Prime Force drones in this scale, it's impossible to complete him.
Note: Although American importers shouldn't have too much trouble, those living in parts of the world using higher strength A/C current outlets should be warned NOT to simply plug the A/C adaptor in with only a socket adapter, as this will overload the device and fry it completely. Buying the appropriate kind of universal A/C adaptor and a stepup/stepdown transformer (the electrical kind) with advice from an electronics specialty shop is recommended.
In Transformers Ongoing, which takes place two years after the events of All Hail Megatron, the Autobots would remain on Earth as Prime feared the return of the Decepticons. Unfortunately, this led to the capture of a number of Autobots by Skywatch, including Windcharger, Gears and Prowl. Hot Rod was reformatted into a Pagani Zonda. Already frustrated, he lashed out at Optimus, accusing him of keeping the Autobots on the planet because he needed someone to protect and demanding he be allowed lead a team to rescue Prowl. An admonished Prime agreed to a covert operation, but with Ironhide along to watch over things. While Prowl was freed, it turned out the humans had used him as bait to draw the other Autobots in. However Wheeljack's new shielding protected Hot Rod's team from Skywatch's inhibitors and in a panic the soldiers switched to lethal measures. Hot Rod was targeted and Ironhide was killed protecting him. An enraged Hot Rod turned on the humans, damaging their equipment and machinery.
If Optimus ordered his troops to come to the humans’ aid, they found themselves taking part in a struggle in futility, as the entire town was near-weightless, meaning they couldn’t put the humans anywhere where they wouldn’t float away once more. Then, the Autobots heard Decepticons approaching. Helpless in their current position, the Autobots were quickly eradicated by the Decepticons.