Originally, Optimus Prime was created as the leader of the Autobots, though it is unknown how he was created or his origins. In a possible future, Megatron was threatened by the existence of the Aerialbots and had Shockwave build a time machine to send them back in time to get rid of them. However, he only ended up changing history for the better. In the revised timeline, Optimus Prime began his life as a robot named Orion Pax, a mostly defenseless dock worker during the Golden Age of Cybertron nine million years ago, with a girlfriend named Ariel and a best friend named Dion. During this time, a new breed of robot with new flight capabilities appeared on the planet that Orion idolized. When Megatron, the leader of the new group of robots, approached him with inquiries about using one of the dock warehouses, Orion was swayed by Megatron. Both Orion and Ariel were severely wounded when Megatron and his forces attacked in order to claim the energy stored there. Searching for someone to help them, the time-displaced Aerialbots took Orion and Ariel to the ancient Autobot, Alpha Trion, who used them as the first subjects for the new reconstruction process he had developed involving rebuilding the frail Autobot frames into more battle-hardy configurations.
A (technically) unreleased "black" redeco of Henkei Voyager-sized Convoy using the same plastic layout and paint mask but replacing red and blue with black, silver with gray, leaving the previously vacuum-metalized grill/fender area unpainted, changing the blue tint for the windows to red and changing all silver and yellow paint operations to blue. The red Autobot insignias would have remained unchanged.
Created with the specific purpose of producing an Optimus Prime as accurate to the original animated series as possible, the toy is exceptionally poseable, partially constructed from die-cast metal, features rubber tires and vacuum-metalized plastic and is heavily detailed, with sliding pistons in its joints and working spring-loaded suspension in vehicle mode. The figure also contains multiple non-intrusive gimmicks, such as a moving mouthplate to emulate Prime's traditional method of speech, flip-up communications panels on the forearms depicting images of Bumblebee and Starscream, and an opening, light-up Matrix chamber in the chest, which contains a removable vacuum-metalized Matrix of Leadership that can also be pulled open.
Alerted by Bumblebee of Megatron's latest plan, Optimus Prime led the mission on Wheeljack's ship to foil them. He sent Bumblebee on an underwater scouting mission, and took part in a pitched battle that ended in the Decepticon's being defeated. Search for Treasure Under the Sea When Rumble dropped rocks on the Autobots, Optimus used Roller to eliminate most of them. Unfortunately he was forced to surrender when he saw the other Autobots had been captured by the Decepticons, and was placed in a special Autobot prison. Luckily Bumblebee remained free, and after he'd rescued the others, Optimus wrapped Megatron up with an iron bar. Bumblebee to the Rescue!
Following a battle which left him grievously wounded, Optimus Prime's entire structural integrity and circuitry processors were reprogrammed into a sleeker, stronger, combat-ready configuration. With a new giant laser-targeting rocket launcher, ripple-fire missile blaster and armor-piercing discs, Optimus was ready to immobilize Megatron once and for all.
Not long after his resurrection, Optimus was alerted by Monitorus to an attack on Tau-Ursa. Optimus's impending speech was interrupted by a Decepticon attack led by Sixshot. Prime soon discovered that the attack was a diversion, while Galvatron tried out his Nullification Cannon on Cybertron's Energon supplies. Optimus rushed to the Energon plant, but, after clouting Slugslinger so hard that the Decepticon was left dangling over a precipice, Prime's morals forced him to rescue the Decepticon instead of stopping Galvatron from getting away.
Across the assorted continuities of the original Transformers universe, there have been various interpretations of Optimus Prime. One of Prime's most notable characteristics over all continuities his unswaying commitment to leadership by example. The animated series' version of Optimus Prime is depicted as a straightforward, wise, and upbeat battlefield general. Additionally, the animated series' version of Optimus dislikes rap music, putting him at odds with music-loving characters like Blaster and Jazz.
Part of the first wave of the 2015 Robots in Disguise line, Optimus Prime is a smaller-scale toy. He transforms from robot to trailer/truck combination and back! Like every Robots in Disguise Legion Class toy, Optimus features 3mm posts on his hands and arms. Due to the size and simplified transformation, he ends up having 6 wheels instead of 10. His scannable Autobot insignia is on his robot mode left shoulder. However, the badge is completely impossible to scan in vehicle mode.
The TakaraTomy "Autobot Alliance" release of this toy has some significant differences, including casting the red plastic in a darker shade, making his shin and wheel joints grey rather than Hasbro's bronze, painting the Matrix of Leadership silver, and swapping out the dual energon swords for battle hooks, similar to 2010's Leader class Optimus Prime. However, due to the Japanese release retaining the Hasbro packaging, the box still advertises the dual energon swords rather than the hooks. He retains the masked face used for the Hasbro release.