Alternity Convoy was simultaneously available in both this silver coloration, and a deco based on his traditional appearance termed "Vibrant Red" (see below). The figure was later redecoed to create "Convoy Super Black" (representing not Optimus Prime, but Universe Nemesis Prime), Alternity Ultra Magnus, and Alternity Dai Atlas. It was retooled into Transformers GT GT-R Prime, GT-R Saber, GT-R Megatron and GT-R Maximus.
This agenda led the Predacon Megatron to attack the comatose Optimus Prime in stasis on prehistoric Earth, forcing Primal to take Prime's Spark into his body in order to preserve his life while his body underwent repairs. Primal was subsequently mutated into the massive "Optimal Optimus" form, which shared elements of Prime's form, prior to returning the Spark to its rightful place. The Maximals were occupied throughout the remainder of the series protecting Prime and the other Transformers aboard the crashed Ark until the Predacons were defeated. Various monuments to Optimus Prime appeared on Cybertron in Beast Machines, one of them a holographic statue in Iacon that Megatron took control of in order to trick Primal.
At some point, Rodimus learned that the world known simply as Planet Q was under threat from the world-devourer, Unicron, but arrived too late to prevent its destruction. Years later, however, he encountered the planet's surviving ruler, Alpha Q, and sided with him in his plan to use Energon to regenerate and restore all the worlds consumed by Unicron – a plan that put him in opposition with Optimus Prime when they encountered each other once again. When Megatron succeeded in reactivating Unicron, however, Rodimus relented and submitted himself and his troops to Optimus Prime's command, and they were integrated into the main Autobot force as the battles continued to defend Alpha Q's successfully-regenerated worlds against the Decepticons.

Optimus Primal, leader of the Maximal faction in the Beast Wars animated series and toy line, is not Optimus Prime. Primal is one of the Maximal descendants of the Autobots, who took the name to honor Optimus Prime. The same applies to the Megatron of this era. Before the animated series began, Hasbro envisioned Prime and Megatron as their beast counterparts, but once the animated series began, this had already been changed. Design elements, such as Prime's mouth-plate slitted to add an actual mouth for the animated series and the first mini-comic that came packaged with the toys, suggest this. Nonetheless, Optimus Prime and Megatron were a major reason the Beast Wars began.


Originally, Optimus Prime was created as the leader of the Autobots, though it is unknown how he was created or his origins. In a possible future, Megatron was threatened by the existence of the Aerialbots and had Shockwave build a time machine to send them back in time to get rid of them. However, he only ended up changing history for the better. In the revised timeline, Optimus Prime began his life as a robot named Orion Pax, a mostly defenseless dock worker during the Golden Age of Cybertron nine million years ago, with a girlfriend named Ariel and a best friend named Dion. During this time, a new breed of robot with new flight capabilities appeared on the planet that Orion idolized. When Megatron, the leader of the new group of robots, approached him with inquiries about using one of the dock warehouses, Orion was swayed by Megatron. Both Orion and Ariel were severely wounded when Megatron and his forces attacked in order to claim the energy stored there. Searching for someone to help them, the time-displaced Aerialbots took Orion and Ariel to the ancient Autobot, Alpha Trion, who used them as the first subjects for the new reconstruction process he had developed involving rebuilding the frail Autobot frames into more battle-hardy configurations.
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