In Transformers: The Last Knight, he is brother-in-arms with Bumblebee who he fought alongside with to foil the Nazi Party in World War 2. He speaks with a thick French accent due to being stuck in France for so long, much to his own embarrassment. He wields a special gun which can briefly slow down time. He is also the only Transformer who is shown to speak in vehicle mode. 
He comes with the Star Saber, which has a 5mm handle and a post on its hilt, as well as two spring-loaded, missile-firing "Ionic Pulse Launchers" (which are called "Eaglefire Missile Launchers" in the instructions). The launchers each feature a 5mm post, have flip open prods, and can peg onto the ports near the wheels on his robot back, giving him a pair of back-mounted rocket engines.
After weeks of Decepticon silence, Optimus Prime decided to get proactive and sent out patrols all over the planet to try and flush the villains out. None of the patrols reported back with any success, however, and Prime realized that there was one area of the planet that his troops had deliberately ignored: the mysterious "Nebulos Triangle". Quickswitch, Scoop and Quickmix were sent into the Triangle, and they did indeed discover and foil the Decepticon plot going on in the area. Decepticons Underground
Brian Colin, Peter Smyth, Mark Buffington, Josh H., Propelstalz, Mary Crocker, Wayne Moulton Jr, Mitch Gross, Jim Valko, Andrew Brown, Lucy Ravitch, Nicholas Duresky, Heather Hofshi, John Kemp, Thorsten Karge, Natasha Dzurny, Keith Ives, Benjamin Chan, Neal Bhatnagar, Justin Farr, Regan Lee, Elaine, Nate Fugal, Stephanie Tennison, Jon Rasmussen, Ryan Pulis, Owen Duffy, Colman Reilly, Anthony, Mark Richman, Alexis Ohanian, Steve, Greg “TVsEgon” Skinner, Andy Saavedra, Daniel, Willie Raymond Taylor III, Chad Ingham, Irene Christian, Clinton Richmond, Jamey Stegmaier, James Allenspach, John Howell, Leif Terry, Tiago Pereira, Nathan Heath, Grinidon, Roman, Berserker Hew, Clark Stacey, Ben Harkins, Kayvaan Ghassemieh, Travis B., Justin Myers, Magna Nordgard, Jim Griffin, Jeff, David Smith, Matthew Titelbaum, Dennis Hitzeman, Daniel Lieske, Michael Jantze, Ruddy, Tom Damico, Kelson, Michelangelo Grigni
It took two further years, but this figure was eventually released as part of the Japanese counterpart to the Alternators line (Binaltech) in 2008. Binaltech Convoy features some additional paint details, like his silver pelvis, additional patterns on his shoulders, and red paint on his engine-gun, and like all Binaltech figures, he also includes die-cast metal pieces. Unlike previous Binaltech releases, a bio-card was not included.
Optimus and his Autobots were aboard the Ark, on the run from the Decepticons. With him, Optimus had what Megatron was forever seeking; the AllSpark! With the ship taking heavy damage from the Autobots' pursuers, Optimus ordered Jazz to take them into an emergency quantum jump. Jazz informed Prime that the jump would take longer to initiate than the time the Decepticons would need to board their vessel. As such, Optimus entrusted the AllSpark to Bumblebee while he and the rest of the bridge crew steeled themselves to hold off their enemies. Thanks to Starscream breaking rank in the Decepticons' formation, Optimus was able to hold back Megatron, and with Heatwave's help, repel them from the ship just as it initiated the quantum jump. However, at that moment the AllSpark tumbled off-ship, falling into the quantum tunnel and seemingly being lost for good... Angry Birds Transformers #1
Released as part of the Allspark Power refresh of the movie line, First Strike Optimus Prime is a extensive redeco of Voyager class Optimus Prime in a colour scheme based upon Generation One and Classics Optimus Prime. The included toy bio describes this coloring as Prime paying tribute to the markings he wore as a young soldier when the war began. Due to the transformation scheme, this figure actually ends up being less like its G1 counterpart in robot mode, a sacrifice made for the slavish redeco of the vehicle mode. You can't have your cake and eat it too, folks.
Hot Rod appears in All Hail Megatron , while traveling through space by himself, he found himself under attack by the Torment, a Decepticon warship crewed by Bludgeon and the Pretender Monsters. He was shot down, but managed to transmit a distress call to the Autobot starship Trion, which came to his rescue. After the Decepticons were chased off, Hot Rod joined the crew of the Trion. When the Decepticons' galaxy-wide assault hit, the Trion was shot down over Cybertron. Hot Rod sent out a distress signal and was ecstatic when he ran into some other Autobots. Hot Rod was taken back to base and relayed the story of how he and his other crew-mates from the Trion arrived on Cybertron, having been diverted there by Decepticon ships and forced to crash land. Once on the surface, Hot Rod sent out his distress signal and was so sure someone heard it. Kup arrived with the rest of the Trion's crew at that point, just in time to tell Hot Rod of course no one heard his message. After the Autobots narrowly avoided the Insecticon swarm and Sunstreaker sacrificed himself, Hot Rod jumped to Drift's defense when he was being insulted by Bumblebee. When the Autobots were later rescued by Omega Supreme, Hot Rod was glad that he sent out the distress call. He was then ferried to Earth with the other Autobots aboard Omega.
Part of the third wave of Prime Cyberverse Commander Class toys, Nightwatch Optimus Prime is a redeco of Series 2 Cyberverse Optimus Prime, using the faceplate-less head sculpt that is gang-molded with the original head. His blue plastic is now gray, his smokestacks and thighs are left unpainted, he loses the Autobot insignia on his right forearm/door, and he gains yellow paint highlights for his eyes and lights, along with various other minor deco differences. Instead of the original release's cannons, he now comes with a pair of swords, which each have a pair of 3mm posts for wielding/mounting.
This redeco of the Ginrai tooling of the Powermaster Optimus Prime figure (released concurrently with the 2001 reissue of the Ginrai figure) was a Toys R Us Japan exclusive black repaint, made to represent the original Optimus Prime equipped for a special mission. All of his red and blue plastics were changed to black, his clear blue windows were now tinted red, making his overall colour scheme very reminiscent of the now-familiar 'Black Convoy' color schemes.
Hot Rod would appear in the UK Marvel comic issue #234, "Prime's Rib!" This story is set in the near future, 1995, where Optimus Prime, Jazz and Hot Rod introduced the latest Autobot, Arcee, to the human feminists. She was met with displeasure by the humans, being called a token female and disliked for her pink color. They were then attacked by Shockwave, Fangry, Horri-Bull and Squeezeplay, who thought the Autobots would be unveiling a new weapon. The Autobots fought off the Decepticons, who escaped, but nothing seemed to please the human feminists.
In shock over the news, Optimus could do little more than scream in agony as Megatron and Starscream made their escape. After regaining his senses, the Autobot leader resumed contact with Ratchet, relaying the order that all Autobots were to return to base. There, Optimus tried his best to calm his panicking subordinates. Though he assured them that power reserves would allow their civilization to continue functioning for some time, they remained jittery. Optimus then declared that he would spearhead a counter-invasion of the Underground. Still, the Autobots remained uncertain, with Brawn pointing out that the Underground was not only vast and dangerous, but mostly unmapped as well. Taking full responsibility for SARA's abduction onto himself, Optimus suggested that he venture to the Underground alone and track SARA's energy signature. Hot Rodimus, however, was unwilling to allow his leader to take such a dangerous course of action, and volunteered to join him. The rest of the Autobots, finding their courage at last, followed suit. Moved by their devotion, Optimus accepted. He also thanked Hot Rodimus for calling out his foolishness, as Optimus now recognized that he would have made an even greater mistake by diving headlong into a doomed venture. Brawn approached Optimus next, insisting he join the group. Optimus at first refused, as Brawn had gotten badly injured during the invasion, but was convinced by the minibot's resolve. Once they were ready, Optimus issued the order to his Autobots... ROLL OUT! Resolve
In the movie, he starts out as a 1976 Camaro before modernizing himself. But the 2007 version (the fifth generation of the model) was a concept car-- you couldn’t even buy a new Camaro until 2009. They later switched to the production version for the 2009 sequel Revenge of the Fallen and he was yet another updated model for Dark of the Moon in 2011 (the SS model). In Age of Extinction, he appears both as a 1967 model and another new model of the current version. Finally, back in June, Bay tweeted that in next year’s The Last Knight, Bee would be a custom-built 2016 Camaro. Over the first two years of the relaunch, Chevy sold an astounding 140,000 Camaros and saw a 10% spike in the yellow model, thanks largely to our friend Bumblebee.
Take note that the leg construction (thigh parts with legs consists of rear wheels/fifth wheel hitch & engine compartment/flip out legs combination) can become loose after a long time of play due to being used with transformation. Also, the legs did not seem to securely connect due to the hitch's shallow holes & the front half of the legs' small pegs. Also, attaching weapons with 3mm posts on the Battle Cannons' holes can cause some stress marks.
As with other earlier Cyber Series figures, Optimus Prime had a rather strange release cycle. At first, he only retailed exclusively in Asia and Latin America, with the closest to a U.S. release being in limited quantities from online retailer BigBadToyStore.com. He did, however, end up being available in the Universal Studios Theme Park, and also later as a Walgreens exclusive.
Megatron is captured by the Autobot Targetmasters in the episode "Do Over". When Optimus learns Megatron has been captured, he rushes to be the one who kills his old enemy, despite the closing launch window of the Ark. When confronted by Optimus Prime, Megatron attempts to use the Autobot Targetmaster Nightstick to shoot the Autobot leader, but Nightstick takes control of Megatron's arm. Megatron then tricks Optimus into using Megatron's own sword to cut his hand off, freeing him from Nightstick. Optimus fails to kill his foe and misses the launching of the Ark, which has been taken over by Rodimus. He then takes out his anger on Blurr.
The TakaraTomy "Autobot Alliance" release of this toy has some significant differences, including casting the red plastic in a darker shade, making his shin and wheel joints grey rather than Hasbro's bronze, painting the Matrix of Leadership silver, and swapping out the dual energon swords for battle hooks, similar to 2010's Leader class Optimus Prime. However, due to the Japanese release retaining the Hasbro packaging, the box still advertises the dual energon swords rather than the hooks. He retains the masked face used for the Hasbro release.