This Japanese series, told through radio plays using the Transformers: Alternators Optimus Prime toy mold, branches off from the original animated series. It is set in a different universe than both the original G1 and Binaltech continuities. Set in the year 2006, one year after Prime's death in The Transformers: The Movie and four years before the third season, it sees Prime's corpse covertly transported to Japan by the Earth Defense Command, a government organization. The convoy transporting his body was attacked by a group of female commandos, led by Marissa Faireborn, who had known Prime years ago as a child. Believing they had secured Prime's body, the commandos were taken by surprise by EDC "Kiss Player" operative Ringo and her Autrooper mechanoid partner, who killed them all except Marissa. Marissa went to Prime's body just as an Autrooper began to fuse with it and, reflecting on her childhood memories of Prime, she gave his faceplate a final kiss. This initiated a transformation where Marissa and Prime were fused together and Prime was reborn with a new body, capable of transforming into a Dodge Ram SRT-10.
An undocumented aspect of his transformation is that his gun, which splits in two halves and secrets inside his kibble when transforming (much like Rattrap's) has two handle pegs on opposite sides, so that it can be held either at its far end or halfway along its length. The former position makes the weapon look (charitably) more like his traditional ion blaster—or Ultra Magnus' rifle. Additionally, the gun halves can be plugged together in two different ways, theoretically allowing you to create an infinitely long weapon (or at least one of ridiculous proportions).
The Autobots continued to guard humanity in secret for many years, until one day in the 21st century when Ratchet was captured by a human group known as Skylogic Systems. Upon investigating the organization, Optimus Prime and Bumblebee came into contact with the Lone Gunmen, three humans who were also on Skylogic's tail for their role in creating a dangerous virus. Learning from them that Ratchet's DNA was part of the concoction, the Autobots joined forces with the humans and helped track down David Simmons, the man responsible for releasing the virus. One interrogation later and they had the address to the Skylogic base, though Prime's size prevented him from easily infiltrating the building, so he left the rescue mission up to Bumblebee and the Gunmen. Once Ratchet was saved, the Autobots parted company with their new human friends in an attempt to lure away any Skylogic pursuers. The X-Files: Conspiracy: The Transformers
The TakaraTomy version of Premium Series Optimus Prime has a considerably different deco from Hasbro's release. Similar to the unique chrome Optimus Prime, Optimus Prime Battle Mode features vacuum-metallized parts for nearly all of the silver truck parts, for a greater resemblance to the polished chrome on the real prop trucks. Unfortunately, as his feet are the front fenders and grill, this will likely lead to chipping. In robot mode, he retains several decos that were removed from Hasbro's Premium Optimus Prime for cost reasons, such as the blue and gold decos on his thighs. Finally, the flip-out energy sword is coloured gold to reflect its "powered up" appearance.
If Optimus led his troops back to the Decepticons’ poison lab, the convoy happened upon the Decepticons hauling barrels of poison out of the poison lab. Windcharger and Ironhide suggested two different plans to Optimus: Windcharger wanted to use his magnetic powers to cause a confusion, while Ironhide wanted to simply go in for a full frontal assault.
The TakaraTomy version, part of the first wave of Movie Advanced Series toys, is dubbed "Classic Optimus Prime", and opts for a deco closer to Prime's previous on-screen appearances, omitting the blue paint on his feet and hands to reveal the gray plastic underneath, and adding new paint details all over. Instead of painting it in clear light blue, his light-piping eyes is neutered with opaque light blue paint. In terms of quality control, at least one sample has missing it's silver paint on the head's left cheek.
The TakaraTomy "Autobot Alliance" release of this toy has some significant differences, including casting the red plastic in a darker shade, making his shin and wheel joints grey rather than Hasbro's bronze, painting the Matrix of Leadership silver, and swapping out the dual energon swords for battle hooks, similar to 2010's Leader class Optimus Prime. However, due to the Japanese release retaining the Hasbro packaging, the box still advertises the dual energon swords rather than the hooks. He retains the masked face used for the Hasbro release.